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For nearly 30 years, biologists have documented a striking pattern of intra-species genetic divergence on the Baja California peninsula in dozens of disparate species. Evolutionary theory predicts that when such a pattern is shared among species the cause is extrinsic (e.g., environmental, climatic, physiographic, geological). The leading hypothesis within biological literature has been that genetic divergence was facilitated by flooding across the central peninsula by a seaway between ~3-1 Ma, resulting in separation of northern and southern populations. However, new detailed geologic mapping from the Baja GeoGenomics consortium reveals evidence for continuous terrestrial environments during the last ~30 Myr in a ≥40-km-wide ~E-W region of the central peninsula that straddles the modern-day crest, conclusively refuting the seaway hypothesis. Through integration of tectonic, volcanic, and sedimentological evidence with genomic (DNA) and gene expression (RNA) data for plants and animals, we are developing a new working model for Earth-life evolution on the peninsula over the last ~5 Myr. In this model, rift-related uplift drives the growth and dissection of topography, causing increased microenvironmental heterogeneity that populations differentially adapted to in the north and south. This is evidenced by widespread, statistically significant niche divergence in populations between northern and southern Baja in 21 studied taxa. This pattern is supported by strong differences in gene expression in northern and southern populations of two lizard species, particularly in genes relating to metabolism, which may indicate different diet or energy requirements between the regions. Habitats in the north and south then shifted due to glacial and interglacial periods, indicated by hindcasting the estimated niche conditions of those 21 taxa. With ongoing analyses, we expect to find genomic signatures of differential natural selection and adaptation within these species due in part to monsoon-driven rainfall differences. The significance of this work is twofold: it demonstrates the importance of incorporating geological data into evolutionary hypotheses and it cautions how mis-assigning cause-effect relationships in individual Earth-life systems can bias our fundamental understanding of how Earth processes shape biological evolution writ large. 

Volcanic rocks of the Sierra San Francisco (SSF), in northern Baja California Sur, Mexico, record post-subduction magmatism related to slab melting and slab window opening. The range is composed of andesitic and dacitic domes, mafic lavas, and volcaniclastic deposits (debris and block-and-ash-flow, lahar, and fluvial) that constitute the proximal to distal facies of a volcanic field with local eruptive ages that postdate the regional transition from subduction to transtension. Lowest observed volcanic units consist of interbedded and hydrothermally altered mafic lavas, tuff breccias, and andesite/dacite domes. These are overlain by volcaniclastic units and dacite domes that erupted between ~11-10 Ma. Volcaniclastic deposits comprise a section up to 800 m thick, locally flank and dip radially away from domes, and are likely associated with dome collapse. These deposits are unconformably overlain by a series of ~5.5-4.5 Ma Mg-enriched basaltic andesites (bajaites) that typically erupted along NNW-trending normal faults. Low interbedded mafic lavas are chemically similar to syn-subduction lavas (>15 Ma) SE of the SSF, suggesting a typical subduction supraslab mantle source during waning, late Miocene Farallon plate subduction. ~11-10 Ma dacite domes and debris flow blocks display an adakitic geochemical signature, implying an origin involving late Miocene foundering and melting of the edges of the subducted Farallon plate during the opening of a slab window after the 12.3 Ma transition from subduction to transtension. Adakitic rocks of the SSF and the Santa Clara volcanic field 60 km to the SW may constrain the E-W extent of the slab window. The ~5.5-4.5 Ma bajaites display enriched REE and trace element patterns, potentially resulting from the rise of enriched subslab mantle through the slab window and interaction with supraslab mantle, previously metasomatized by slab melts. Thermal pulses associated with Gulf of California rifting may have provided the heat to generate Mg-rich magmas which ascended along rift-related faults, precluding significant crustal contamination or fractionation, and allowing magmas to retain their primitive character. Further analysis will elucidate the timing of slab window development and the post-subduction mantle processes that drove the chemical evolution of SSF magmas. 

The Sierra San Francisco (SSF) is a Neogene volcanic range along the topographic crest of the Baja California peninsula in northern Baja California Sur, Mexico. The SSF is ~55 km long (NW-SE) and ~30 km wide and its highest peaks exceed 1500 m elevation. The SSF has a long history of volcanism and has been eroded by deep, rugged, radially-draining canyons. The development of SSF topography is intimately associated with the volcanic evolution of the range. The SSF is a large and complex dacitic adakite dome complex largely built of a thick, up to 800 m, stratigraphic succession of dacitic tuff breccias with minor interbedded basaltic andesite lavas. These deposits overlie rare exposures of aeolian sandstone of unknown age. The tuff breccias represent block-and-ash-flows and lahars generated from steep-sided peleean dacite and andesite domes, with three radiometric dates of 11-10 Ma. This intermediate sequence is unconformably capped by widespread bajaite mafic lavas, 5.5-4.5 Ma. SSF topography evolved dramatically since the late Miocene: 1) From 11-10 Ma, adakite domes erupted across the central SSF, locally along NNW faults. Thick sequences of bedded tuff breccias accumulated around the domes and are radially inclined away from source domes. The duration of this volcanism is unknown. 2) From 10-5 Ma, deep erosion of the pyroclastic strata formed a range-wide radial drainage network, with channel depths of up to 130 m or more. 3) From 5.5-4.5 Ma, voluminous bajaite lavas from cinder cones and dike vents flooded the top of the range and flowed down the radial drainages with flow distances up to 12 km. Vents are strongly aligned along steep NNW normal faults. 4) After 4.5 Ma, erosion removed interfluves of tuff breccia not armored by younger mafic lavas. Today, the long, steep-sided, lava-capped ridges are inverted topographically. At Santa Martha, an area in the central SSF with the highest concentration of domes, hydrothermal alteration of the volcanic deposits during and after the dome volcanism caused severe material weakening and slope failure within the volcanic center. The area is now a distinctive erosional basin, partly filled with clay-rich landslide deposits. Comparable volcanic history and topographic development are likely to have occurred in a dome field of similar age and size at Santa Agueda, 60 km SE of Santa Martha. 

ABSTRACT New paleoseismic trenching indicates late Quaternary oblique right-lateral slip on the Leech River fault, southern Vancouver Island, Canada, and constrains permanent forearc deformation in northern Cascadia. A south-to-north reduction in northward Global Navigation Satellite System velocities and seismicity across the Olympic Mountains, Strait of Juan de Fuca (JDF), and the southern Strait of Georgia, has been used as evidence for permanent north–south crustal shortening via thrust faulting between a northward migrating southern forearc and rigid northern backstop in southwestern Canada. However, previous paleoseismic studies indicating late Quaternary oblique right-lateral slip on west-northwest-striking forearc faults north of the Olympic Mountains and in the southern Strait of Georgia are more consistent with forearc deformation models that invoke oroclinal bending and(or) westward extrusion of the Olympic Mountains. To help evaluate strain further north across the Strait of JDF, we present the results from two new paleoseismic trenches excavated across the Leech River fault. In the easternmost Good Hope trench, we document a vertical fault zone and a broad anticline deforming glacial till. Comparison of till clast orientations in faulted and undeformed glacial till shows evidence for postdeposition faulted till clast rotation, indicating strike-slip shear. The orientation of opening mode fissuring during surface rupture is consistent with right-lateral slip and the published regional SHmax directions. Vertical separation and the formation of scarp-derived colluvium along one fault also indicate a dip-slip component. Radiocarbon charcoal dating within offset glacial till and scarp-derived colluvium suggest a single surface rupturing earthquake at 9.4±3.4  ka. The oblique right-lateral slip sense inferred in the Good Hope trench is consistent with slip kinematics observed on other regional west-northwest-striking faults and indicates that these structures do not accommodate significant north–south shortening via thrust faulting.